E3b, what is this 2002? There is a 22,000 year gap between E-M35 and E-M78. E-M78 in the Horn derives from the pre-Neolithic Maghreb and is not from the Horn of Africa. It is a closed case with the finding of the Taforalt samples who emerged precisely around the TMRCA of E-M78. You can't get any closer than that with an ancient lineage.
I never said E-M78 is from the Horn of Africa, IMO it is Egyptian and would have been introduced into the Horn during the Neolithic. I said E-M215/E-M35 originated in the Horn of Africa but that population itself would have been genetically North African.
This model below makes the most sense IMO.
There is also no evidence that the E1b1b1 lineages found in Berbers or the one in Natufians came recently from the Horn of Africa around the time of the mergence of proto-Afro-Asiatic (~15K YA). Their lineages could have been in Northern Africa for over 30,000 years and have zero ties to Omotic lineages.
E1b1b1 lineages in Taforalt/Natufians would have come from Egypt, but that population would have come from the Horn of Africa, who in turn, would have come from MSA Egypt, there were bidirectional migrations between the Horn and Egypt. The core ancestry of that population would have been native to North Africa.
There is Egyptian ancestry in both the Natufians and Taforalt, they also share Omotic/Hadza/Mota related ancestry, so whilst these Egyptians took refuge in the Horn they mixed with those people and spread Mota related ancestry in the Levant and Taforalt. I don't think that population would have spoke Afro Asiatic but maybe pre-proto Afro-Asiatic, something ancestral to Afro Asiatic. I predict proto-Afro asiatic speakers would be predominantly Natufian/Taforalt related and/or rich in Basal Eurasian with some Mota/Hadza related ancestry and maybe some Dinka. BTW, idk much about Afro Asiatic from a linguistic standpoint, I think Omotic is Afro Asiatic purely because it fits with genetics and it seems to be the general consensus among scholars, I'm just addressing the points you raised regarding genetics.
I never see clear evidence of an earlier migration of Afrasians in Ethiopia predating Cushites and linking it to the rest of the AA world.
Mota carried little to no Neanderthal, but can be modelled as part Natufian related, this could represent a pre-neolithic migration from Egypt. If it was during the neolithic than they would show some appreciable neanderthal ancestry, but they don't.
"Helwan lunates are extremely rare, and given their particular hafting technique, one may wonder if these were not borrowed from other, older or contemporary assemblages. Interestingly,
in southern Sinai,
the assemblages of the Abu Madi I site (Bar-Yosef 1985), dated to ca. 9,600-8,300 cal BC, contain el-Khiam and tanged points, as well as small rods (bipolar retouched, narrow, double pointed microliths) and a few
Helwan lunates. It seems that a site that lies some 150-250 km south of any Natufian sites including those on both sides of the Jordan Rift valley retained an old tradition. Abu Madi I is also far away from the original localities at Helwan, where a couple of dozens of Helwan lunates were found in the detailed survey carried out by F. Debono in the 1930s (Schmidt 1996 and references therein)."
"To this we should also add the undated context of obsidian Helwan lunates in an assemblage retrieved in
Dahlak island (Eritrea) in the Red Sea some 1,800 km south of the Nile delta (Blanc 1952). By comparison to studied shell middens with lunates, the dates at Dahlak may range from ca.
6,800 to 6,000 cal BC (Bar Yosef Mayer and Beyin 2009)."
This attests to migration/contact between the Horn and Egypt, just before the Neolithic, which definitely predates Cushites in the Horn and could represent a source of ancestry for Mota/Omotic types.